Nitrococcus mobilis: Difference between revisions

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When free floating in the ocean, Nitrococcus mobilis exists as a single or diploid cell. If the cells can form a biofilm then several hundred will form aggregates on a slime layer.  
When free floating in the ocean, Nitrococcus mobilis exists as a single or diploid cell. If the cells can form a biofilm then several hundred will form aggregates on a slime layer.  


The cell is a cocci with a diameter of 1.5 to 1.7 micrometers after division. Nitrococcus mobilis is the only nitrifier that is coccus. A 3-4 micrometer long flagella aids in mobility when the cell is free floating. If apart of a biofilm, the flagella is lost and the only mode of movement is by rolling which often lacks a direction. Nitrococcus mobilis is gram negative as it has an outer cell membrane surrounding a multilayered cell wall. In the cytosol of Nitrococcus mobilis is a unique cytomembrane system. This tubuler system is continuous with the plasma membrane. The plasma membrane will invagenate, at what appears to be randomly distributed, to form the tubes. The tubes themselves are 6-8 nm in diameter and are formed by a double tracked cytomembrane. Electron microscopy has shown that the tubes do have contents in them. Other distinct structures in the cytosol are inclusion bodies
The cell is a cocci with a diameter of 1.5 to 1.7 micrometers after division. Nitrococcus mobilis is the only nitrifier that is coccus. A 3-4 micrometer long flagella aids in mobility when the cell is free floating. If apart of a biofilm, the flagella is lost and the only mode of movement is by rolling which often lacks a direction. Nitrococcus mobilis is gram negative as it has an outer cell membrane surrounding a multilayered cell wall. In the cytosol of Nitrococcus mobilis is a unique cytomembrane system. This tubuler system is continuous with the plasma membrane. The plasma membrane will invagenate, at what appears to be randomly distributed, to form the tubes. The tubes themselves are 6-8 nm in diameter and are formed by a double tracked cytomembrane. Electron microscopy has shown that the tubes do have contents in them. Other distinct structures in the cytosol are inclusion bodies. These are aggregates of insoluble proteins. It is unclear if the insolubility is due to misfolding or if the proteins are in different states. The particles are surrounded by a single layered membrane. The most prominent inclusion bodies are hexagonal in shape and have a diameter of approximately 100 nm. A single cell can contain as many as 200 of these particles. The cells also have glycogen storage areas and poly-beta-hydroxybutyrate granules.


==Ecology and Pathogenesis==
==Ecology and Pathogenesis==

Revision as of 21:13, 23 March 2010

Classification

Domain: Bacteria Phylum: Proteobacteria Class: Gammaproteobacteria Order: Chromatiales Family: Ectothiorhodospiraceae

Species

NCBI: Taxonomy

Nitrococcus mobilis

Description and Significance

Describe the appearance, habitat, etc. of the organism, and why you think it is important.

Genome Structure

Describe the size and content of the genome. How many chromosomes? Circular or linear? Other interesting features? What is known about its sequence?


Cell Structure, Metabolism and Life Cycle

Interesting features of cell structure; how it gains energy; what important molecules it produces.

Nitrococcus mobilis is an obligate chemoautotroph. The energy required for metabolism comes from chemical reactions of reducing nitrogen from nitrite to nitrate. They are aerobic as oxygen is the final electron acceptor. Inorganic carbon is its carbon source, making it an autotroph.

When free floating in the ocean, Nitrococcus mobilis exists as a single or diploid cell. If the cells can form a biofilm then several hundred will form aggregates on a slime layer.

The cell is a cocci with a diameter of 1.5 to 1.7 micrometers after division. Nitrococcus mobilis is the only nitrifier that is coccus. A 3-4 micrometer long flagella aids in mobility when the cell is free floating. If apart of a biofilm, the flagella is lost and the only mode of movement is by rolling which often lacks a direction. Nitrococcus mobilis is gram negative as it has an outer cell membrane surrounding a multilayered cell wall. In the cytosol of Nitrococcus mobilis is a unique cytomembrane system. This tubuler system is continuous with the plasma membrane. The plasma membrane will invagenate, at what appears to be randomly distributed, to form the tubes. The tubes themselves are 6-8 nm in diameter and are formed by a double tracked cytomembrane. Electron microscopy has shown that the tubes do have contents in them. Other distinct structures in the cytosol are inclusion bodies. These are aggregates of insoluble proteins. It is unclear if the insolubility is due to misfolding or if the proteins are in different states. The particles are surrounded by a single layered membrane. The most prominent inclusion bodies are hexagonal in shape and have a diameter of approximately 100 nm. A single cell can contain as many as 200 of these particles. The cells also have glycogen storage areas and poly-beta-hydroxybutyrate granules.

Ecology and Pathogenesis

Habitat; symbiosis; biogeochemical significance; contributions to environment.
If relevant, how does this organism cause disease? Human, animal, plant hosts? Virulence factors, as well as patient symptoms.

References

[Sample reference] Takai, K., Sugai, A., Itoh, T., and Horikoshi, K. "Palaeococcus ferrophilus gen. nov., sp. nov., a barophilic, hyperthermophilic archaeon from a deep-sea hydrothermal vent chimney". International Journal of Systematic and Evolutionary Microbiology. 2000. Volume 50. p. 489-500.

Author

Page authored by Hanna Miller and Toni Miskovich, students of Prof. Jay Lennon at Michigan State University.